Senior Lecturer
Institute of Aquaculture Pathfoot Building, University of Stirling, Stirling, FK9 4LA
Dr Darren Green MA PhD DIC I am interested the ecology of disease, how diseases spread from place to place, why particular populations are more at risk than others, and how we can control and prevent disease outbreaks. Oct 2007 - Present University of Stirling, Institute of Aquaculture Lecturer in Aquatic Health Modelling. My research involves a variety of numerical approaches to tackling problems of aquatic animal health discussed more in the "projects" section below. May 2004 - Aug 2007 University of Oxford, Department of ZoologyPost-doctoral research assistant. Research centred on the modelling and epidemiology of disease. The approaches include network models of disease spread, deterministic differential-equation-based models, and individual-based microsimulation. Aspects of this research were featured twice on Farming Today and recently were reported in the Daily Telegraph online.May 2000 - May 2004 University of Edinburgh, School of Geosciences (form. IERM) Post-doctoral research associate. Work as part of a DEFRA LINK research programme. My research was centred on the production of a simulation model of animal growth response to nutrition and the environment. The animal concerned was the domestic pig. The model was then tested against growth trial data from dissection and monitoring of live animals through imaging by video camera (visual image analysis, VIA). October 1996 - January 2000 Imperial College, University of London.PhD Biological Sciences October 1993 - June 1996 Clare College, University of Cambridge BA Natural Sciences (Zoology) Outside interests: A player of the piano and viola, I also sing bass or tenor, whichever's in the shortest supply. I also enjoy hill-walking, running, cycling, and mint KitKats.
My research is focussed on the epidemiology of fish diseases, however I have a continuing interest in terrestrial farming systems.
ROBUST-SMOLT Impact of Early Life History in Freshwater Recirculation Aquaculture Systems on A. Salmon Robustness and Susceptibility to Disease at Sea
PI: Professor Herve Migaud
Funded by: Biotechnology and Biological Sciences Research Council
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Introduction to Mathematical Modelling for the environmental and biological sciences
PI: Dr Andrew Hoyle
Funded by: Natural Environment Research Council
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Introduction to mathematical modelling for the environmental and biological sciences
PI: Professor Rachel Norman
Funded by: Natural Environment Research Council
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Introduction to mathematical modelling for the environmental and biological sciences
PI: Professor Rachel Norman
Funded by: Natural Environment Research Council
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Development of epidemiological models for the spread and control of bluetongue virus
PI: Dr Darren Green
Funded by: The Institute for Animal Health
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Book Chapter
The Global Approach: Aquaculture and National Action Plans for Antimicrobial Resistance
Desbois A & Green D (2025) The Global Approach: Aquaculture and National Action Plans for Antimicrobial Resistance. In: Elumalai P & Lakshmi S (eds.) Antimicrobial Resistance in Aquaculture and Aquatic Environments. London, pp. 385-412. https://doi.org/10.1007/978-981-97-7320-6_16
Article
Green D (2025) A new web application for determining sample size in freedom-from-disease testing with imperfect tests. Preventive Veterinary Medicine, 235, Art. No.: 106397. https://doi.org/10.1016/j.prevetmed.2024.106397
Other
Aquaculture requires special consideration in National Action Plans for Antimicrobial Resistance
Desbois AP, Brunton LA, Henriksson PJG, Luthman O, Troell M & Green DM (2025) Aquaculture requires special consideration in National Action Plans for Antimicrobial Resistance. Science of The Total Environment, 958, Art. No.: 177785. https://doi.org/10.1016/j.scitotenv.2024.177785
Article
The birthday problem: repeated sampling of animal populations and ethics of experimental design
Green DM & Mitchell CG (2024) The birthday problem: repeated sampling of animal populations and ethics of experimental design. animal, 18 (11), Art. No.: 101352. https://doi.org/10.1016/j.animal.2024.101352
Article
Corrochano-Fraile A, Carboni S, Green DM, Taggart JB, Adams TP, Aleynik D & Bekaert M (2024) Estimating blue mussel (Mytilus edulis) connectivity and settlement capacity in mid-latitude fjord regions. Communications Biology, 7, Art. No.: 166. https://doi.org/10.1038/s42003-023-05498-3
Article
When and why direct transmission models can be used for environmentally persistent pathogens
Benson L, Davidson RS, Green DM, Hoyle A, Hutchings MR & Marion G (2021) When and why direct transmission models can be used for environmentally persistent pathogens. PLOS Computational Biology, 17 (12), Art. No.: e1009652. https://doi.org/10.1371/journal.pcbi.1009652
Article
Metselaar M, Thompson KD, Paley R, Green DM, Verner-Jeffreys D, Feist S & Adams A (2020) Investigating the involvement of a Midichloria -like organism (MLO) in red mark syndrome in rainbow trout Oncorhynchus mykiss. Aquaculture, 528, Art. No.: 735485. https://doi.org/10.1016/j.aquaculture.2020.735485
Article
Network structure and risk-based surveillance algorithms for live shrimp movements in Thailand
Saleetid N & Green DM (2019) Network structure and risk-based surveillance algorithms for live shrimp movements in Thailand. Transboundary and Emerging Diseases, 66 (6), pp. 2450-2461. https://doi.org/10.1111/tbed.13303
Article
The Contact Structure of Great Britain's Salmon and Trout Aquaculture Industry
Jones AE, Munro LA, Green DM, Morgan KL, Murray AG, Norman R, Salama NKG, Ryder D, Taylor NGH, Thrush MA, Wallace IS & Sharkey KJ (2019) The Contact Structure of Great Britain's Salmon and Trout Aquaculture Industry. Epidemics, 28, Art. No.: 100342. https://doi.org/10.1016/j.epidem.2019.05.001
Article
Orton R, Deason M, Bessell PR, Green D, Kao RR & Salvador LCM (2018) Identifying genotype specific elevated-risk areas and associated herd risk factors for bovine tuberculosis spread in British cattle. Epidemics, 24, pp. 34-42. https://doi.org/10.1016/j.epidem.2018.02.004
Article
Sumner T, Orton R, Green D, Kao RR & Gubbins S (2017) Quantifying the roles of host movement and vector dispersal in the transmission of vector-borne diseases of livestock. PLoS Computational Biology, 13 (4), Art. No.: e1005470. https://doi.org/10.1371/journal.pcbi.1005470
Article
Time-to-response toxicity analysis as a method for drug susceptibility assessment in salmon lice
Carmona-Antoñanzas G, Humble JL, Carmichael SN, Heumann J, Christie HRL, Green DM, Bassett DI, Bron JE & Sturm A (2016) Time-to-response toxicity analysis as a method for drug susceptibility assessment in salmon lice. Aquaculture, 464, pp. 570-575. https://doi.org/10.1016/j.aquaculture.2016.08.007
Article
Sirimanapong W, Thompson KD, Ooi EL, Bekaert M, Collet B, Taggart J, Bron J, Green D, Shinn A, Adams A & Leaver M (2015) The effects of feeding β-glucan to Pangasianodon hypophthalmus on immune gene expression and resistance to Edwardsiella ictaluri. Fish and Shellfish Immunology, 47 (1), pp. 595-605. https://doi.org/10.1016/j.fsi.2015.09.042
Article
Seasonality in live fish movements and its effects on epidemic dynamics
Werkman M, Murray AG, Munro LA, Turnbull J & Green D (2014) Seasonality in live fish movements and its effects on epidemic dynamics. Aquaculture, 418-419, pp. 72-78. https://doi.org/10.1016/j.aquaculture.2013.10.001
Article
Oidtmann B, LaPatra SE, Verner-Jeffreys DW, Pond M, Peeler EJ, Noguera PA, Bruno DW, St-Hilaire S, Schubiger CB, Snekvik K, Crumlish M, Green D, Metselaar M, Rodger HD, Schmidt-Posthaus H, Galeotti M & Feist SW (2013) Differential characterization of emerging skin diseases of rainbow trout - a standardized approach to capturing disease characteristics and development of case definitions. Journal of Fish Diseases, 36 (11), pp. 921-937. https://doi.org/10.1111/jfd.12086
Conference Paper (unpublished)
Sirimanapong W, Ooi EL, Thompson KD, Bekaert M, Collet B, Taggart JB, Bron JE, Green DM, Adams A & Leaver MJ (2013) Effect of B-glucan on immune response genes of Pangasianodon hypophthalmus and protection against Ewardsiella ictalurI. First Conference of the International Society of Fish and Shellfish Immunology, 25.06.2013-28.06.2013.
Article
Factors affecting variation in mortality of marine Atlantic salmon Salmo salar in Scotland
Soares S, Murray AG, Crumlish M, Turnbull J & Green D (2013) Factors affecting variation in mortality of marine Atlantic salmon Salmo salar in Scotland. Diseases of Aquatic Organisms, 103 (2), pp. 101-109. https://doi.org/10.3354/dao02562
Article
Soares S, Murray AG, Crumlish M, Turnbull J & Green D (2012) Evaluating abnormal mortality as an indicator of disease presence in the Atlantic salmon industry using the receiver operating characteristic (ROC). Aquaculture, 370-371, pp. 136-143. https://doi.org/10.1016/j.aquaculture.2012.10.004
Article
The Impact of Escaped Farmed Atlantic Salmon (Salmo salar L.) on Catch Statistics in Scotland
Green D, Penman D, Migaud H, Bron J, Taggart J & McAndrew B (2012) The Impact of Escaped Farmed Atlantic Salmon (Salmo salar L.) on Catch Statistics in Scotland. PLoS ONE, 7 (9), p. e43560. https://doi.org/10.1371/journal.pone.0043560
Article
From Markovian to pairwise epidemic models and the performance of moment closure approximations
Taylor M, Simon P, Green D, House T & Kiss IZ (2012) From Markovian to pairwise epidemic models and the performance of moment closure approximations. Journal of Mathematical Biology, 64 (6), pp. 1021-1042. https://doi.org/10.1007/s00285-011-0443-3
Article
Review of climate change impacts on marine aquaculture in the UK and Ireland
Callaway R, Shinn A, Grenfell SE, Bron J, Burnell G, Cook EE, Crumlish M, Culloty S, Davidson K, Ellis RP, Flynn KJ, Fox C, Green D, Hays GC, Hughes AD, Johnston E, Lowe CD, Lupatsch I, Malham SK, Mendzil AF, Nickell T, Pickerell T, Rowley AF, Stanley MS, Tocher DR, Turnbull J, Webb G, Wootton E & Shields RJ (2012) Review of climate change impacts on marine aquaculture in the UK and Ireland. Aquatic Conservation: Marine and Freshwater Ecosystems, 22 (3), pp. 389-421. https://doi.org/10.1002/aqc.2247
Article
The potential for targeted surveillance of live fish movements in Scotland
Green D, Werkman M & Munro LA (2012) The potential for targeted surveillance of live fish movements in Scotland. Journal of Fish Diseases, 35 (1), pp. 29-37. https://doi.org/10.1111/j.1365-2761.2011.01321.x
Article
Seasonality and heterogeneity of live fish movements in Scottish fish farms
Werkman M, Green D, Munro LA, Murray AG & Turnbull J (2011) Seasonality and heterogeneity of live fish movements in Scottish fish farms. Diseases of Aquatic Organisms, 96 (1), pp. 69-82. https://doi.org/10.3354/dao02382
Article
Applied epidemiology with examples from UK aquaculture
Turnbull J, Berrill I, Green D, Kaye R, Morris D, Murray AG, del Pozo Gonzalez J & Shinn A (2011) Applied epidemiology with examples from UK aquaculture. Aquaculture Research, 42 (Supplement s1), pp. 21-27. https://doi.org/10.1111/j.1365-2109.2010.02667.x
Article
Tools to study trends in community structure: application to fish and livestock trading networks
Green D, Werkman M, Munro LA, Kao RR, Kiss IZ & Danon L (2011) Tools to study trends in community structure: application to fish and livestock trading networks. Preventive Veterinary Medicine, 99 (2-4), pp. 225-228. http://www.sciencedirect.com/science/journal/01675877; https://doi.org/10.1016/j.prevetmed.2011.01.008
Article
A baseline method for benchmarking mortality losses in Atlantic salmon (Salmo salar) production
Soares S, Green D, Turnbull J, Crumlish M & Murray AG (2011) A baseline method for benchmarking mortality losses in Atlantic salmon (Salmo salar) production. Aquaculture, 314 (1-4), pp. 7-12. https://doi.org/10.1016/j.aquaculture.2011.01.029
Article
Werkman M, Green D, Murray AG & Turnbull J (2011) The effectiveness of fallowing strategies in disease control in salmon aquaculture assessed with an SIS model. Preventive Veterinary Medicine, 98 (1), pp. 64-73. http://www.sciencedirect.com/science/journal/01675877; https://doi.org/10.1016/j.prevetmed.2010.10.004
Article
Mapping of AF1 transactivation domains in duplicated rainbow trout glucocorticoid receptors
Sturm A, Bron J, Green D & Bury NR (2010) Mapping of AF1 transactivation domains in duplicated rainbow trout glucocorticoid receptors. Journal of Molecular Endocrinology, 45 (6), pp. 391-404. https://doi.org/10.1677/JME-09-0152
Article
Large-scale properties of clustered networks: implications for disease dynamics
Green D & Kiss IZ (2010) Large-scale properties of clustered networks: implications for disease dynamics. Journal of Biological Dynamics, 4 (5), pp. 431-445. https://doi.org/10.1080/17513758.2010.487158
Article
del Pozo Gonzalez J, Crumlish M, Ferguson H, Green D & Turnbull J (2010) A prospective longitudinal study of "Candidatus arthromitus"-associated rainbow trout gastroenteritis in the UK. Preventive Veterinary Medicine, 94 (3-4), pp. 289-300. https://doi.org/10.1016/j.prevetmed.2010.02.001
Article
A strategic model for epidemic control in aquaculture
Green D (2010) A strategic model for epidemic control in aquaculture. Preventive Veterinary Medicine, 94 (1/2), pp. 119-127. http://www.sciencedirect.com/science/journal/01675877; https://doi.org/10.1016/j.prevetmed.2009.12.004
Article
Semigroup analysis of structured parasite populations
Farkas JZ, Green D & Hinow P (2010) Semigroup analysis of structured parasite populations. Mathematical Modelling of Natural Phenomena, 5 (3), pp. 94-114. http://www.mmnp-journal.org/action/displayJournal?jid=MNP; https://doi.org/10.1051/mmnp/20105307
Article
Small- and large-scale network structure of live fish movements in Scotland
Green D, Gregory A & Munro LA (2009) Small- and large-scale network structure of live fish movements in Scotland. Preventive Veterinary Medicine, 91 (41001), p. 261; 269. http://www.sciencedirect.com/science/journal/01675877; https://doi.org/10.1016/j.prevetmed.2009.05.031
Article
Coevolution of dispersal in a parasitoid-host system
Green D (2009) Coevolution of dispersal in a parasitoid-host system. Population Ecology, 51 (2), pp. 253-260. https://doi.org/10.1007/s10144-008-0131-3
Commentary
Comment on "Properties of highly clustered networks"
Kiss IZ & Green D (2008) Comment on "Properties of highly clustered networks". Commentary on: MEJ Newman, 'Properties of Highly Clustered Networks', Phys. Rev. E 68, 026121 (2003). Physical Review E, 78 (4), Art. No.: 048101. https://doi.org/10.1103/PhysRevE.78.048101
Article
Kiss IZ, Green D & Kao RR (2008) The effect of network mixing patterns on epidemic dynamics and the efficacy of disease contact tracing. Journal of the Royal Society Interface, 5 (24), pp. 791-799. https://doi.org/10.1098/rsif.2007.1272
Article
Estimates for local and movement-based transmission of bovine tuberculosis in British cattle
Green D, Kiss IZ, Mitchell AP & Kao RR (2008) Estimates for local and movement-based transmission of bovine tuberculosis in British cattle. Proceedings of the Royal Society B: Biological Sciences, 275 (1638), pp. 1001-1005. http://rspb.royalsocietypublishing.org/content/275/1638/1001.abstract; https://doi.org/10.1098/rspb.2007.1601
Article
Demographic risk factors for classical and atypical scrapie in Great Britain
Green D, del Rio Vilas VJ, Birch CPD, Johnson J, Kiss IZ, McCarthy ND & Kao RR (2007) Demographic risk factors for classical and atypical scrapie in Great Britain. Journal of General Virology, 88 (12), pp. 3486-3492. https://doi.org/10.1099/vir.0.83225-0
Article
Kao RR, Green D, Johnson J & Kiss IZ (2007) Disease dynamics over very different time-scales: foot-and-mouth disease and scrapie on the network of livestock movements in the UK. Journal of the Royal Society Interface, 4 (16), pp. 907-916. https://doi.org/10.1098/rsif.2007.1129
Article
Data quality of the Cattle Tracing System in Great Britain
Green D & Kao RR (2007) Data quality of the Cattle Tracing System in Great Britain. Veterinary Record, 161 (13), pp. 439-443. https://doi.org/10.1136/vr.161.13.439
Article
Real-time control of pig growth through an integrated management system
Parsons DJ, Green D, Schofield CP & Whittemore CT (2007) Real-time control of pig growth through an integrated management system. Biosystems Engineering, 96 (2), pp. 257-266. https://doi.org/10.1016/j.biosystemseng.2006.10.013
Article
Parasite strain coexistence in a heterogeneous host population
Green D, Kiss IZ & Kao RR (2006) Parasite strain coexistence in a heterogeneous host population. Oikos, 115 (3), pp. 495-503. https://doi.org/10.1111/j.2006.0030-1299.15155.x
Article
Modelling the initial spread of foot-and-mouth disease through animal movements
Green D, Kiss IZ & Kao RR (2006) Modelling the initial spread of foot-and-mouth disease through animal movements. Proceedings of the Royal Society B: Biological Sciences, 273 (1602), pp. 2729-2735. https://doi.org/10.1098/rspb.2006.3648
Article
Kiss IZ, Green D & Kao RR (2006) The network of sheep movements within Great Britain: network properties and their implications for infectious disease spread. Journal of the Royal Society Interface, 3 (10), pp. 669-677. https://doi.org/10.1098/rsif.2006.0129
Article
The effect of contact heterogeneity and multiple routes of transmission on final epidemic size
Kiss IZ, Green D & Kao RR (2006) The effect of contact heterogeneity and multiple routes of transmission on final epidemic size. Mathematical Biosciences, 203 (1), pp. 124-136. https://doi.org/10.1016/j.mbs.2006.03.002
Article
Demographic structure and pathogen dynamics on the network of livestock movements in Great Britain
Kao RR, Danon L, Green D & Kiss IZ (2006) Demographic structure and pathogen dynamics on the network of livestock movements in Great Britain. Proceedings of the Royal Society B: Biological Sciences, 273 (1597), pp. 1999-2007. https://doi.org/10.1098/rspb.2006.3505
Article
Parameterization of individual-based models: Comparisons with deterministic mean-field models
Green D, Kiss IZ & Kao RR (2006) Parameterization of individual-based models: Comparisons with deterministic mean-field models. Journal of Theoretical Biology, 239 (3), pp. 289-297. https://doi.org/10.1016/j.jtbi.2005.07.018
Article
Infectious disease control using contact tracing in random and scale-free networks
Kiss IZ, Green D & Kao RR (2006) Infectious disease control using contact tracing in random and scale-free networks. Journal of the Royal Society Interface, 3 (6), pp. 55-62. https://doi.org/10.1098/rsif.2005.0079
Article
Disease contact tracing in random and clustered networks
Kiss IZ, Green D & Kao RR (2005) Disease contact tracing in random and clustered networks. Proceedings of the Royal Society B: Biological Sciences, 272 (1570), pp. 1407-1414. https://doi.org/10.1098/rspb.2005.3092
Article
Calibration and sensitivity analysis of a model of the growing pig for weight gain and composition
Green D & Whittemore CT (2005) Calibration and sensitivity analysis of a model of the growing pig for weight gain and composition. Agricultural Systems, 84 (3), pp. 279-295. https://doi.org/10.1016/j.agsy.2004.06.017
Article
The relationship between body dimensions of living pigs and their carcass composition
Doeschl-Wilson AB, Green D, Fisher AV, Carroll SM, Schofield CP & Whittemore CT (2005) The relationship between body dimensions of living pigs and their carcass composition. Meat Science, 70 (2), pp. 229-240. https://doi.org/10.1016/j.meatsci.2005.01.010
Article
The relationship between the body shape of living pigs and their carcass morphology and composition
Doeschl-Wilson AB, Green D, Whittemore CT, Schofield CP, Fisher AV & Knap PW (2004) The relationship between the body shape of living pigs and their carcass morphology and composition. Animal Science, 79, pp. 73-83.
Article
White RP, Schofield CP, Green D, Parsons DJ & Whittemore CT (2004) The effectiveness of a visual image analysis (VIA) system for monitoring the performance of growing/finishing pigs. Animal Science, 78, pp. 409-418.
Article
Whittemore CT, Green D, Wood JD, Fisher AV & Schofield CP (2003) Physical and chemical composition of the carcass of three different types of pigs grown from 25 to 115 kg live weight. Animal Science, 77, pp. 235-245.
Article
Growth of carcass components and its relation with conformation in pigs of three types
Fisher AV, Green D, Whittemore CT, Wood JD & Schofield CP (2003) Growth of carcass components and its relation with conformation in pigs of three types. Meat Science, 65 (1), pp. 639-650. https://doi.org/10.1016/S0309-1740%2802%2900266-8
Article
Green D, Brotherstone S, Schofield CP & Whittemore CT (2003) Food intake and live growth performance of pigs measured automatically and continuously from 25 to 115 kg live weight. Journal of the Science of Food and Agriculture, 83 (11), pp. 1150-1155. https://doi.org/10.1002/jsfa.1519
Article
Green D & Whittemore CT (2003) Architecture of a harmonized model of the growing pig for the determination of dietary net energy and protein requirements and of excretions into the environment (IMS Pig). Animal Science, 77, pp. 113-130.
Article
The determination of beginning- and end-of-period live weights of growing pigs
Schofield CP, Whittemore CT, Green D & Pascual MD (2002) The determination of beginning- and end-of-period live weights of growing pigs. Journal of the Science of Food and Agriculture, 82 (14), pp. 1672-1675. https://doi.org/10.1002/jsfa.1247
Article
The description of the rate of protein and lipid growth in pigs in relation to live weight
Whittemore CT & Green D (2002) The description of the rate of protein and lipid growth in pigs in relation to live weight. Journal of Agricultural Science, 138 (4), pp. 415-423. https://doi.org/10.1017/S0021859602002186
Article
Technical review of the energy and protein requirements of growing pigs: protein
Whittemore CT, Green D & Knap PW (2001) Technical review of the energy and protein requirements of growing pigs: protein. Animal Science, 73, pp. 363-373.
Article
Technical review of the energy and protein requirements of growing pigs: energy
Whittemore CT, Green D & Knap PW (2001) Technical review of the energy and protein requirements of growing pigs: energy. Animal Science, 73, pp. 199-215.
Article
Technical review of the energy and protein requirements of growing pigs: food intake
Whittemore CT, Green D & Knap PW (2001) Technical review of the energy and protein requirements of growing pigs: food intake. Animal Science, 73, pp. 3-17.
Article
Evolutionary interactions between Drosophila melanogaster and its parasitoid Asobara tabida
Green D, Kraaijeveld AR & Godfray HCJ (2000) Evolutionary interactions between Drosophila melanogaster and its parasitoid Asobara tabida. Heredity, 85 (5), pp. 450-458. https://doi.org/10.1046/j.1365-2540.2000.00788.x
